FACULTY

R. Stephen Howard Professor of Biology Ph.D, Indiana University, 1994 Office: 615-898-2044 Fax: 615-898-5093 E-mail: rshoward@mtsu.edu

"There is a general place in your brain, I think, reserved for "melancholy of relationships past." It grows and prospers as life progresses, forcing you finally, against your grain, to listen to country music."

- Kary B. Mullis, from his acceptance speech for the 1993 Nobel Prize in Chemistry.

Representative papers:

S. G. Johnson and R. S. Howard (2007). Contrasting patterns of synonymous and nonsynonymous sequence evolution in asexual and sexual freshwater snail lineages. Evolution

R. S. Howard and C. M. Lively (2004). Good vs Complementary genes for parasite resistance and the evolution of female mate choice. BMC Evolutionary Biology 4:48

J. F. Fontanari, A. Colato, and R. S. Howard (2003). Mutation accumulation in growing asexual lineages. Physical Review Letters 91 (21) 218101

R. S. Howard and C. M. Lively (2003). Opposites attract? Mate choice for parasite evasion and the evolutionary stability of sex. Journal of Evolutionary Biology (16) 681-689

R. S. Howard and C. M. Lively (2002). The Ratchet and the Red Queen: the maintenance of sex in parasites. Journal of Evolutionary Biology (15) 648-656

Some older papers:

Theoretical Population Biology:
Nature:
Evolution:
Philosophical Transactions of the Royal Society: